Daisy Wilder prefers the company of horses to people, bare feet and salt water to high heels and society parties. Then, in the dizzying aftermath of the attack on Pearl Harbor, Daisy enlists in a top secret program, replacing male soldiers in a war zone for the first time. Under fear of imminent invasion, the WARDs guide pilots into blacked-out airstrips and track unidentified planes across Pacific skies. But not everyone thinks the women are up to the job, and the new recruits must rise above and work side by side despite the resistance and heartache they meet along the way.

In the summer of 1940, ambitious young American journalist Ruby Sutton gets her big break: the chance to report on the European war as a staff writer for Picture Weekly newsmagazine in London. She jumps at the chance, for it's an opportunity not only to prove herself, but also to start fresh in a city and country that know nothing of her humble origins. But life in besieged Britain tests Ruby in ways she never imagined.

In a different world, Evie Milton would have accepted her fate, married an aristocrat and become the doyenne of one of England’s finest estates, just like her mother. In a different world, Rose Wiley would have married her fiancé, David, established a modest homestead and brought up a brood of babies, just like her mother. But this isn’t a different world, and these women are not their mothers. Rose dreams of a life filled with more than family and duty to her husband—a life of purpose—and Evie dreams of a life far away from her rarefied existence.

Twenty-five years after the end of the war, Noah Ainsworth is still preoccupied with those perilous, exhilarating years as a British SOE operative in France. A head injury sustained on his final operation has caused frustrating gaps in his memory—in particular about the agent who saved his life during that mission gone wrong, whose real name he never knew, nor whether she even survived the war. Moved by her father’s frustration, Noah’s daughter Charlotte begins a search for answers that resurrects the stories of Chloe and Fleur, the code names for two otherwise ordinary women.

1941. Audrey Coltrane has always wanted to fly. It's why she implored her father to teach her at the little airfield back home in Texas. It’s why she signed up to train military pilots in Hawaii when the war in Europe began. And it’s why she insists she is not interested in any dream-derailing romantic involvements, even with the disarming Lieutenant James Hart, who fast becomes a friend as treasured as the women with whom she flies. Then one fateful day, she gets caught in the air over Pearl Harbor just as the bombs begin to fall, and suddenly, nowhere feels safe.