Ruby Lennox begins narrating her life at the moment of conception, and from there takes us on a whirlwind tour of the 20th century as seen through the eyes of an English girl determined to learn about her family and its secrets.

Living on her family’s gorgeous lakeside estate in Cornwall, England, Alice Edevane is a bright, clever, inquisitive, innocent, and precociously talented fourteen-year-old who loves to write stories. But the mysteries she pens are no match for the one her family is about to endure ...One midsummer’s eve, after a beautiful party drawing hundreds of guests to the estate has ended, the Edevanes discover that their youngest son, Theo, has vanished without a trace.

As England enters World War II's dark early days, spirited music professor Primrose Trent, recently arrived to the village of Chilbury, emboldens the women of the town to defy the Vicar's stuffy edict to shutter the church's choir in the absence of men and instead carry on singing. Resurrecting themselves as The Chilbury Ladies' Choir, the women of this small village soon use their joint song to lift up themselves and the community as the war tears through their lives.

Born into poverty, brutalised and ignored by her family, it is only when she is married off to a nobleman that her life undergoes a transformation, as her husband recognises in Rosamund a magic she does not know she possesses but which affects almost all who encounter her. Presiding over a luxurious chocolate house where men go to be seen, exchange news and indulge in the sweet and heady drink to which they have become addicted, Rosamund feels blessed. But disaster strikes, and Rosamund stands on the brink of losing all she has worked so hard to achieve.

After Englishman John Blackthorne is lost at sea, he awakens in a place few Europeans know of and even fewer have seen—Nippon. Thrust into the closed society that is seventeenth-century Japan, a land where the line between life and death is razor-thin, Blackthorne must negotiate not only a foreign people, with unknown customs and language, but also his own definitions of morality, truth, and freedom. As internal political strife and a clash of cultures lead to seemingly inevitable conflict, Blackthorne’s loyalty and strength of character are tested by both passion and loss.