When Viney Walker’s long-absent father arrives in the 19th-century utopian community of Rugby, TN, he begs her to return with him to the Great Smoky Mountains. Viney’s sister, Lizzie, urges her to go because a new setting will help Viney heal from a broken engagement. Viney acquiesces, and in her new home, she meets her Walker cousins, including handsome and brawny James. The couple’s romance angers the White Caps, a vigilante group that whips lewd women, and they warn Viney to mend her ways. Seeking revenge and the freedom to love James, Viney joins a counter-vigilante group.

On October 28, 1940, Mussolini provides Greek Prime Minister Ioannis Metaxas with an ultimatum - either allow Axis forces to occupy their country or face war - and Greece’s response is swift: “Oxi!” they say. “No!” In a small village nestled against the radiant waters of the Aegean Sea, Alexei, the son of a local fisherman, and his best friend Costa - who were both born on the same night eighteen years earlier and have been like brothers ever since - leave their homes, like all the other young men in their village and throughout Greece, to bravely fight for their country.

Two New York girls, Maria Mezza and Alice Doe, wake up one November morning in 1906. Their lives are about to change forever. Maria is heading to her first job in a shirtwaist factory wishing she was heading to the country side. Alice is leaving New York to find a new family in Illinois country side. Three years later, the paths of the two girls will converge once more. Gone is the innocence of youth yet both girls are desperately hoping their lives will improve. Maria is caught up in the Shirtwaist Strike while Alice is on the run for her life.`

England, 1941. After three nights of relentless bombing from German aircraft, trained nurse Jo Brooks is told to report to the basement theater of Mill Road Hospital. She goes with a heavy heart, not wanting to leave behind her best friend Moira, who is desperately soothing new mothers on the maternity ward. As Jo arrives safely underground, the ward takes a direct hit. 

England, 1943: Home is no longer safe for eight-year-old twins Molly and Jacob. Night after night, wailing bombs and screeching planes skim the rooftops overhead. They cradle each other, shivering in terror, not knowing if they'll live to see dawn. Their mother, Martha, has no choice but to evacuate them to the safety of the countryside. At the train station, Martha bites back tears as she says good-bye to her precious children. Knowing she might never see them again, she gives Jacob a letter, pressing the envelope into his hands and telling him to only read it if they are in danger.

The attack on Pearl Harbor changes everything for Lana Hitchcock. Arriving home on the Big Island too late to reconcile with her estranged father, she is left alone to untangle the clues of his legacy, which lead to a secret property tucked away in the remote rain forest of Kilauea volcano. When the government starts taking away her neighbors as suspected sympathizers, Lana shelters two young German girls, a Japanese fisherman, and his son. As tensions escalate, they are forced into hiding - only to discover the hideaway house is not what they expected.

Sarah Jennings knows there’s one place she can go to find some peace and quiet during her difficult divorce. But arriving at her beloved father’s home in the countryside, she finds him unwell and hunched over boxes of files, studying the records from Cedar Hall, the crumbling orphanage in town. He says that hidden behind the wrought iron gates and overgrown ivy are secrets about their family, and he asks for her help.  

Liverpool, 1939. Bella Rogers thought her life was just starting; she can’t wait to get married to her childhood sweetheart, Bobby. But when war is declared and Bobby is called up, his letters stop, leaving Bella heartbroken and alone. Then her family is torn apart by tragedy when her angelic 5-year-old sister dies from a terrible illness. They can barely afford a funeral, but the church is packed with the entire community, devastated for their loss. Grieving for her sister and lost love, Bella finds a job at the Bryant and May factory, making matches.

Bridget Collins is in dire straits - she needs to get out of New York, fast. With two young siblings under her wing, her options are limited. Her priest sends her as an outplacement agent on the orphan trains that run from New York to out west. With almost 40 orphans under her care, she's relieved that fellow and more experienced outplacement agent Carl Watson is there to guide her. But Carl is dealing with his own trauma and finds it difficult to handle the pain the orphans are dealing with.

It’s a miracle when Katherine’s baby boy is born healthy. But his twin sister doesn’t survive, and when Katherine is told she can’t have any more children, the loss is almost too much for her to bear. Katherine always saw herself having a big family: She remembers how the loneliness of being an only child used to overwhelm her, and she is desperate to adopt a sibling for her son. But her husband, Davis, won’t agree. He worries that Katherine will struggle when the new baby arrives.

Kathleen Green longs for a child of her own. Her work with the orphan trains only serve to heighten her need. When tragedy strikes, will she get the child she always wanted? Will the cost be worth it? 

When Ava’s partner Will is diagnosed with a terminal brain tumour, the doctors give Will one chance to survive - an operation which means he will lose his recent memory. Ava begs him to take the chance, sure that she can cope with Will forgetting her. After all, they have something very special to live for. But they are also keeping a heart-breaking secret, and if Will loses his memory, Ava will have to carry that secret alone. Can they rebuild their love from scratch, or will their secrets and past come between them?

September 1940. As the bombs of the Blitz fall on London, Ruby and Stevie are falling in love. United by a shocking experience when they were evacuees, Ruby believes that she understands Stevie like nobody else can. But then Stevie is sent abroad into danger, and as Ruby waits desperately for letters with foreign stamps that never come, she begins to fear that he is lost forever. And when Stevie does return, he has changed, and Ruby faces a difficult choice. 

It's 1911, and Lily's plans for her factory are not progressing as fast as she would like. Maria, her sister Rosa, and friend Leonie are still working at the Triangle Shirtwaist Factory. Maria and Conrad have agreed it's pointless finding new jobs when they will soon be joining Lily's factory. Rosa can think of nothing but her upcoming marriage, and Leonie worries how she is going to look after her four siblings when their mother falls very ill.  

1945: Two sisters give birth to two little girls on the same night, huddled under blankets, deep in the black woods that surround their village. They hold their babies close as footsteps approach. If they make even the slightest sound, the German soldiers will find them....