Florence, the 1550s. Lucrezia, third daughter of the grand duke, is comfortable with her obscure place in the palazzo: free to wonder at its treasures, observe its clandestine workings, and to devote herself to her own artistic pursuits. But when her older sister dies on the eve of her wedding to the ruler of Ferrara, Moderna and Regio, Lucrezia is thrust unwittingly into the limelight: the duke is quick to request her hand in marriage, and her father just as quick to accept on her behalf. 

When the bohemian, sophisticated Innes Kent turns up by chance on her doorstep, Lexie Sinclair realises she cannot wait any longer for her life to begin, and leaves for London. There, at the heart of the 1950s Soho art scene, she carves out a new life for herself, with Innes at her side. In the present day, Elina and Ted are reeling from the difficult birth of their first child.

England, 1580. A young Latin tutor - penniless, bullied by a violent father - falls in love with an eccentric young woman: a wild creature who walks her family's estate with a falcon on her shoulder and is known throughout the countryside for her unusual gifts as a healer.

Edinburgh in the 1930s. The Lennox family is having trouble with its youngest daughter. Esme is outspoken, unconventional, and repeatedly embarrasses them in polite society. Even Kitty, Esme's beloved sister, is beginning to lose patience. Something will have to be done.

Tuscany, 1944: As Allied troops advance and bombs fall around deserted villages, a young English soldier, Ulysses Temper, finds himself in the wine cellar of a deserted villa. There, he has a chance encounter with Evelyn Skinner, a middle-aged art historian who has come to Italy to salvage paintings from the ruins and recall long-forgotten memories of her own youth. In each other, Ulysses and Evelyn find a kindred spirit amidst the rubble of war-torn Italy, and set off on a course of events that will shape Ulysses's life for the next four decades.