Thirteen-year-old Lisa has escaped from Nazi Germany on the Kindertransport. She arrives in London unable to speak a word of English, her few belongings crammed into a small suitcase. Among them is one precious photograph of the family she has left behind. Lonely and homesick, she is adopted by a childless couple. But when the Blitz blows her new home apart, she wakes up in a hospital with no memory of who she is or where she came from. The authorities give her a new name and dispatch her to a children's home. With the war raging around her, what will become of Lisa now?

Plymouth, 1941. As sirens blare all around, the Shawbrook family take refuge in a packed shelter. Bombs have already begun to fall through the night sky when they realize their infant son, Freddie, has been forgotten in the rush, left to sleep in his crib. Terrified, Vera, his young mother, races to find him and bring him to safety. The next morning, air-raid warden David Shawbrook returns from his watch to find the shelter pulverized, and his family seemingly all dead. Dirty footprints inside their home betray the looters who have rifled through the house.

Florence, South Carolina 1944: Lieutenant Raymond Sellers bids farewell to his brother Jimmie, the weight of what lies ahead hanging heavy in the air. The brothers have always been inseparable—Raymond, the steady protector, Jimmie, the wild-hearted dreamer. But as Raymond embraces his pregnant wife Evelyn one last time, war is about to test the bonds of family in ways they never imagined. As a summer storm shakes the windows, Evelyn brings her baby into a world at war.

It is two years before the outbreak of World War 1, and the Oakley family are typical of their time. Andrew has been clerk at a City solicitors for twenty years. Alice, his wife, keeps the home fires burning. Their sons plan on becoming skilled apprentices. And Mabel, their daughter, says she would rather die than go into domestic service like her cousin. She is going to go to train as a secretary. Andrew Oakley little imagines, as he sets off for work one February morning, that their lives are about to change in the most dramatic and tragic fashion.

Rita and Rosie Stevens are only nine and five years old when their widowed mother marries a violent bully called Jimmy Randall and has a baby boy by him. Under pressure from her new husband, she is persuaded to send the girls to an orphanage, not knowing that the papers she has signed will entitle them to do what they like with the children. And it is not long before the powers that be decide to send a consignment of orphans to their sister institution in Australia. Among them - without their family's consent or knowledge - are Rita and Rosie.