Four months after the explosion at the Garden, a place where young women known as the Butterflies were kept captive, FBI agents Brandon Eddison, Victor Hanoverian, and Mercedes Ramirez are still entrenched in the aftermath, helping survivors in the process of adjusting to life on the outside. With winter coming to an end, the Butterflies have longer, warmer days of healing ahead.

Eight-year-old Brooklyn Mercer has gone missing. And as accustomed as FBI agents Eliza Sterling and Brandon Eddison are to such harrowing cases, this one has struck a nerve. It marks the anniversary of the disappearance of Eddison’s own little sister. Disturbing, too, is the girl’s resemblance to Eliza - so uncanny they could be mother and daughter. With Eddison’s unsettled past rising again with rage and pain, Eliza is determined to solve this case at any cost. But the closer she looks, the more reluctant she is to divulge to her increasingly shaken partner what she finds.

Near an isolated mansion lies a beautiful garden. In this garden grow luscious flowers, shady trees...and a collection of precious “butterflies” - young women who have been kidnapped and intricately tattooed to resemble their namesakes. Overseeing it all is the Gardener, a brutal, twisted man obsessed with capturing and preserving his lovely specimens.

Janey Markham is thrilled to be moving with her family to Buckingham Crescent, the smartest address in a desirable suburban town. Worried she’ll be excluded by the glossy local mothers, Janey is thrilled when she meets Tanya, the kind of woman she has always looked up to. Tanya takes Janey under her wing and her teenage daughter, Angel, is amazing with Janey’s little boy. As Janey and Tanya grow closer, Janey feels she can finally leave her troubled past behind.  

Stephanie and Patrick are adjusting to life with their colicky twin girls. The babies are a handful, but even as Stephanie struggles with the disorientation of sleep deprivation, there's one thing she's sure of: She has all she ever wanted. Then Erica, a woman from Patrick's past, appears and makes a disturbing accusation. Patrick had always said his first wife's death was an accident, but now Erica claims it was murder. Patrick insists he's innocent, that this is nothing but a blackmail attempt. Still, Erica knows things about Patrick.